Steve Trombulak

Professor in the Department of Biology and Program in Environmental StudiesStephen C. Trombulak
Anderson Professor of Environmental and Biosphere Studies
Middlebury College
Middlebury, Vermont 05753 U.S.A.

B.A. Biology (now, Ecology and Evolutionary Biology), University of California, Los Angeles (1977)
Ph.D. Zoology (now, Biology), University of Washington (1983)

Director, Conservation Biology Laboratory at Middlebury College
Editor, Journal of Natural History Education and Experience

Former Director, Middlebury School of the Environment

Miscellaneous videos related to my professional work:

Faculty Stories: Steve Trombulak
Owl banding (Vermont Outdoor Journal on VPT)
Reclaiming the Soul of Higher Education (TEDx presentation)

Research interests

My professional activities are divided among four fields: ecology, conservation biology, natural history, and environmental science education. I have ongoing research and outreach projects in the following areas:

  • Theoretical concepts related to the protection and restoration of biological integrity
  • Regional conservation issues in the Northern Appalachian/Acadian ecorgion (northeastern U.S. and southeastern Canada)
  • Ecology and conservation of carabid beetles in Vermont
  • Incorporating education on critical thinking and scientific methodology into environmental science curricula
  • Fostering a renaissance in the practice and appreciation of natural history

Classes taught

Environmental Science (ES 112)
Vertebrate Biology (BIOL 202)
Vertebrate Natural History (BIOL 302)
Conservation Biology (BIOL 392)
Senior Seminar in Environmental Studies (ENVS 401)

Recent publications

Baldwin, R.F., S.C. Trombulak, P.B. Leonard, R.F. Noss, J.A. Hilty, H.P. Possingham, L. Scarlett, and M.G. Anderson. 2018. The future of landscape conservation. BioScience: in press.

Fleischner, T.L., R.E. Espinoza, G.A. Gerrish, H.W. Greene, R.W. Kimmerer, E.A. Lacey, S. Pace, J.K. Parrish, H.M. Swain, S.C. Trombulak, S. Weisberg, D. W. Winkler, and L. Zander. 2017. Teaching biology in the field: Importance, challenges, and solutions. BioScience 67: 558-567.

Perkl, R.M., R.F. Baldwin, S.C. Trombulak, and G.R. Smith. 2016. Landscape network congruency at the ecoregion scale: can we expect corridor ‘umbrellas’? Journal of Land Use Science, DOI: 10.1080/1747423X.2016.1142015.

Witt, J., H. Peterson, and S.C. Trombulak. 2016. Case Study: ‘Understanding Place’ at the Middlebury School of the Environment. Journal of Sustainability Education 11: ISSN 2151-7452.

D’Aloia, C.C., C.B. Azodi, S.P. Sheldon, S.C. Trombulak, and W.R. Ardren. 2015. Genetic models reveal historical patterns of sea lamprey population fluctuations within Lake Champlain. PeerJ 3:e1369, DOI 10.7717/peerj.1369.

Klyza, C.M., and S.C. Trombulak. 2015. The Story of Vermont: A natural and cultural history (2nd edition). Hanover, NH: University Press of New England.

Schmitz, O.J., J.J. Lawler, P. Beier, C. Groves, G. Knight, D.A. Boyce Jr., J. Bulluck, K.M. Johnston, M.L. Klein, K. Muller, D.J. Pierce, W.R. Singleton, J.R. Strittholt, D.M. Theobald, S.C. Trombulak, and A. Trainor. 2015. Conserving biodiversity: practical guidance about climate change adaptation approaches in support of land-use planning. Natural Areas Journal 35: 190-203.

Tewksbury, J.J., J.G.T. Anderson, J.D. Bakker, T.J. Billo, P.W. Dunwiddie, M.J. Groom, S.E. Hampton, S.G. Herman, D.J. Levey, N.J. Machnicki, C. Martinez del Rio, M.E. Power, K. Rowell, A.K. Salomon, L. Stacey, S.C. Trombulak, and T.A. Wheeler. 2014. Natural history’s place in science and society. BioScience 64: 300-310.

Halfacre, A.C., J. Byrne, M. Horhota, K. Kransteuber, S.C. Trombulak, B. DeKnight, B. Andersen, and N. Jenks-Jay. 2013. Shaping sustainability at Furman and Middlebury: Emergent and adaptive curricular models. In High Education for Sustainability: Cases, Challenges, and Opportunities from Across the Curriculum (L.F. Johnston, ed.). New York: Routledge.

Rudnick, D.A., S.J. Ryan, P. Beier, S. Cushman, F. Dieffenbach, C.W. Epps, L. Gerber, J. Hartter, J. Jenness, J. Kintsch,  A.M. Merenlender, R.M. Perkl, D.V. Preziosi, and S.C. Trombulak.  2012.  The role of landscape connectivity in planning and implementing conservation and restoration priorities. Issues in Ecology 16: 1-20.

Trombulak, S.C., R.F. Baldwin, J.J. Lawler, J. Hepinstall-Cymerman, and M.G. Anderson. 2012. The Northern Appalachian/Acadian ecoregion, North America.  In Climate and Conservation: Landscape and seascape science, planning, and action (J.A. Hilty, C.C. Chester, and M.S. Cross, eds.).  Washington, D.C.: Island Press.

Al-Abdulrazzak, D., and S.C. Trombulak.  2012. Classifying levels of protection in marine protected areas. Marine Policy 36: 576-582.

Noss, R.F., A.P. Dobson, R. Baldwin, P. Beier, C.R. Davis, D.A. Dellasala, J. Francis, H. Locke, A. Nowak, R. Lopez, C. Reining, S.C. Trombulak, and G. Tabor. 2012. Bolder thinking for conservation. Conservation Biology 26: 1-4.

Chester, C.C., J.A. Hilty, and S.C. Trombulak. 2012. Climate change science, impacts, and opportunities. In Climate and Conservation: Landscape and seascape science, planning, and action (J.A. Hilty, C.C. Chester, and M.S. Cross, eds.).  Washington, D.C.: Island Press.

Trombulak, S.C. 2011. Becoming a neighbor.  In The Way of Natural History (T. Fleischner, ed.) Trinity University Press.

Trombulak, S.C., and R.F. Baldwin, editors. 2010. Landscape-scale Conservation Planning. New York: Springer.

Baldwin, R.F., R. Perkl, S.C. Trombulak, S.C., and W.B. Burwell. 2010. Modeling ecoregional connectivity. In Landscape-scale Conservation Planning (S.C. Trombulak and R.F. Baldwin, eds.) New York: Springer.

Hamilton, L.S., and S.C. Trombulak. 2010. Greater Northern Appalachian bioregion. Pp. 152-160 in Connectivity Conservation Management: A Global Guide (G.L. Worboys, W.L. Francis, M. Lockwood, eds.). Earthscan: London.

Trombulak, S.C., and R.F. Baldwin. 2010. Introduction: Creating a context for landscape-scale conservation planning. In Landscape-scale Conservation Planning (S.C. Trombulak and R.F. Baldwin, eds.) New York: Springer.

Trombulak, S.C., R.F. Baldwin, and G. Woolmer. 2010. The Human Footprint as a conservation planning tool. In Landscape-scale Conservation Planning (S.C. Trombulak and R.F. Baldwin, eds.) New York: Springer.

Trombulak, S.C.. 2010. Assessing irreplaceability in systematic conservation planning. In Landscape-scale Conservation Planning (S.C. Trombulak and R.F. Baldwin, eds.) New York: Springer.

Baldwin, R.F., S.C. Trombulak, and E.D. Baldwin. 2009. Assessing risk of large-scale habitat conversion in lightly settled landscapes. Landscape and Urban Planning. doi: 10.1016/j.landurbplan.2009.01,004.

Richmond, L., and S.C. Trombulak. 2009. Distribution of red-backed salamander (Plethodon cinereus) with respect to cover-object characteristics in the Green Mountains of Vermont. Northeastern Naturalist 16: 13-26.

Trombulak, S.C., M.G. Anderson, R.F. Baldwin, K. Beazley, J.C. Ray, C. Reining, G. Woolmer, C. Bettigole, G. Forbes, and L. Gratton. 2008. The Northern Appalachian/Acadian Ecoregion: Priority Locations for Conservation Action. Two Countries, One Forest / Deux Pays, Une Forêt Special Report 1.

Woolmer, G., S.C. Trombulak, J. Ray, P. Doran, M. Anderson, R. Baldwin, A. Morgan, and E. Sanderson. 2008. Rescaling the Human Footprint: A tool for conservation planning at an ecoregional scale. Landscape and Urban Planning 87: 42-53.

Baldwin, R.F., J.C. Ray, S.C. Trombulak, G. Woolmer. 2007. Relationship between spatial distribution of urban sprawl and species imperilment: response to Brown and Laband. Conservation Biology 21: 546-548.

Baldwin, R.F., S.Trombulak, M. Anderson and G. Woolmer. 2007. Projecting transition probabilities for regular public roads at the ecoregion scale: a Northern Appalachian/Acadian case study. Landscape and Urban Planning 80: 404-411.

Trombulak, S.C. 2007. Wilderness and wild lands in the Northern Appalachian region of North America: An ecological perspective. Pp. 510-514 in Science and stewardship to protect and sustain wilderness values: Eighth World Wilderness Congress symposium (A. Watson, J. Sproull, and L. Dean, eds.). Proceedings RMRS-P-49. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. Fort Collins, CO.

Baldwin, R.F., and S.C. Trombulak. 2007. Losing the dark: a case for a national policy on land conservation. Conservation Biology 21: 1133-1134.

Daly, H.E., B. Czech, D.L. Trauger, W.E. Rees, M. Grover, T. Dobson, S.C. Trombulak. 2007. Are we consuming too much – for what? Conservation Biology 21: 1359-1362.

Trombulak, S.C., and T.L. Fleischner. 2007. Natural history renaissance. Journal of Natural History Education 1:1-4.

Baldwin, R.F., S.C. Trombulak, G. Woolmer, K. Beazley, C. Reining, J. Nordgren, and M. Anderson. 2007. The importance of Maine for ecoregional conservation planning. Maine Policy Review 16: 66-77.

Other activities

Advisory Committee, Vermont Invertebrate Database Alliance

I used to be involved in a lot more, but seriously, being both the Director of Sciences and Director of the Middlebury School of the Environment takes up a lot of time!

plus, some rules to live by

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